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  1. Departments of Ophthalmology & Visual Sciences, and,
  2. Department of Physiology, University of Kentucky, Lexington, Kentucky 40506, USA
  3. Department of Chemistry, The University of North Carolina at Chapel Hill, Chapel Hill, North Carolina 27599-3290, USA
  4. Department of Ophthalmology, Nagoya City University Medical School, Nagoya 467-8601, Japan
  5. Casey Eye Institute, Oregon Health and Science University, Portland, Oregon 97239, USA
  6. Department of Ophthalmology and Visual Sciences, Moran Eye Center,
  7. Program in Human Molecular Biology and Genetics, Eccles Institute of Human Genetics, University of Utah School of Medicine, Salt Lake City, Utah 84132, USA
  8. Department of Neurosurgery, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania 19104, USA
  9. Department of Ophthalmology, Veterans Affairs Salt Lake City Healthcare System, Salt Lake City, Utah 84148, USA
  10. Center for Wound Healing & Tissue Regeneration, University of Illinois at Chicago College of Dentistry, Chicago, Illinois 60612-7211, USA
  11. Laboratory for Molecular Medicine, University of North Carolina at Greensboro, Greensboro, North Carolina 27402-6170, USA
  12. These authors contributed equally to this work.